Robbie Henshaw will partner Bundee Aki in Japan of the World Cup at centre in Irelands Pool A decider against Samoa on Saturday.
The Irish and British Lions power has missed all three of Irelands World Cup matches so much, with picked up a hamstring injury in practice after landing in Japan.
However, that the 26-year-old is fit and ready to fire to some game Ireland should win to guarantee their place at the quarter-finals.
Ireland have gathered the majority of their big guns to take on the assaulting and bodily Samoans, however Jordan Larmour has slotted in at full-back instead of Rob Kearney.
Tadhg Beirne replenishes his Munster colleague Peter OMahony at blindside flanker, together with manager Joe Schmidt eager to ease the workload on the British and Irish Lions back-rower.
While Joey Carbery takes a seat on the seat, with Ireland buoyed to possess play-makers back to full fitness johnny Sexton begins at fly-half center.
Ireland: 15 Jordan Larmour, 14 Keith Earls, 13 Robbie Henshaw,12 Bundee Aki, 11 Jacob Stockdale, 10 Jonathan Sexton, 9 Conor Murray, 1 Cian Healy, 2 Rory Best (c), 3 Tadhg Furlong, 4 Iain Henderson, 5 James Ryan, 6 Tadhg Beirne, seven Josh van der Flier, 8 CJ Stander
Replacements: 23 Andrew Conway, 16 Niall Scannell, 17 Dave Kilcoyne, 18 Andrew Porter, 19 Jean Kleyn 21 Luke McGrath, 22 Joey Carbery
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